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Öğe A new species of Euscorpius Thorell, 1876 (Scorpiones, Euscorpiidae) from Turkey(Pensoft Publishers, 2012) Tropea, Gioele; Yagmur, Ersen Aydin; Koc, Halil; Yesilyurt, Fatih; Rossi, AndreaA new species of the genus Euscorpius Thorell, 1876 is described based on specimens collected from Dilek Peninsula (Davutlar, Aydin) in Turkey. It is characterized by an oligotrichous trichobothrial pattern (Pv= 7, et= 5/6, eb= 4) and small size. Euscorpius (Euscorpius) avcii sp. n. is the first named species of the subgenus Euscorpius from Turkey.Öğe A new species of Zachaeus CL Koch from Turkey (Opiliones, Phalangiidae)(Pensoft Publ, 2015) Kurt, Kemal; Koc, Halil; Yagmur, Ersen AydinA new species of harvestmen, Zachaeus seyyari sp. n. (Opiliones, Phalangiidae), is described and illustrated on the basis of both sexes from Sirnak Province in Turkey. Differences between the new species and related species are indicated. Photographs of its characteristic structures are also provided.Öğe A revised checklist of the Solifugae (Arachnida) of Greece(Kmk Scientific Press Ltd, Moscow State Univ, 2023) Koc, HalilAn overview of Greek solifuges is presented as an annotated checklist of the bibliogra-phy, covering the literature related directly to the solif-uges of Greece with all known localities. Two species are added to the known camel spider fauna for the country, Galeodes araneoides (Pallas, 1772) and G. barbarus Lucas, 1849, which were omitted from the last checklist. Currently, a total of 14 species and one subspecies belonging to four families and eight genera are reported from Greece. Four species, Gluviella rhodi-ensis Caporiacco, 1948, Galeodes hellenicus Roewer, 1934, G. rhodicola Roewer, 1941 and Barrussus fur-cichelis Roewer, 1928 and one genus, Gluviella , are presently known to be endemic. Potential erroneous data and dubious distributional records that may exist in the literature are also identified.Öğe Checklist of the Solifugae (Arachnida) of Iraq(Kmk Scientific Press Ltd, Moscow State Univ, 2023) Koc, HalilA checklist of all taxa of the order Solifugae from Iraq is presented. The checklist is based on the compilation of data gathered from published records up to now. This list comprises 21 species and 3 subspecies belonging to 11 genera distributed in 4 fam-ilies, Daesiidae, Galeodidae, Karschiidae and Rhago-didae. It composes of one endemic genus; Rhagodos-pus, and four endemic species; Galeodes adamsi (Turk, 1947), G. lawrencei Harvey, 2002 (Galeodidae), Rha-godixa kurdistanica (Birula, 1936) and Rhagodospus babylonicus (Birula, 1935) (Rhagodidae). Two previ-ously overlooked species, Gluviopsilla discolor (Krae-pelin, 1899) (Daesiidae) and Galeodes bacillifer Po-cock, 1900 (Galeodidae) are included in the list. Alto-gether, five species erroneously reported from the coun-try are omitted from the checklist; Galeodes bacillifer-oides Roewer, 1934, G. starmuehlneri Roewer, 1952, G. sulfuripes Roewer, 1934, G. taurus (Roewer, 1934) and Gylippus (Gylippus) syriacus (Simon, 1872). This study provides more detailed and comprehensive docu-mentation of solifuges in Iraq.Öğe Comparative cheliceral morphology of the solifuge species Biton zederbaueri (Werner) and Gluviopsilla discolor (Kraepelin) through scanning electron microscopy (Arachnida: Solifugae: Daesiidae)(Max Kasparek Verlag, 2011) Bayram, Abdullah; Yigit, Nazife; Erdek, Melek; Koc, Halil; Sancak, Zafer; Melekoglu, Abdullah; Ocal, Ilkay CorakA detailed comparative morphology of the male chelicerae of Biton zederbaueri (Werner) and Gluviopsilla discolor (Kraepelin) is presented using stereo microscope and scanning electron microscopy (SEM). The morphology of the cheliceral teeth, setae, flagella and some other organs such as the stridulatory organ that are used for description and identification are described and illustrated. The flagella are membranous, but their shapes arc different in both species. The stridulatory organs are similar in both daesiids, being reniform and with seven ridges. On the chelicerae, numerous long-thin, short-thin and short-thick setae are characteristic for both species.Öğe Evolutionary dynamics of rDNA clusters on chromosomes of buthid scorpions (Chelicerata: Arachnida)(Oxford Univ Press, 2020) Stahlavsky, Frantisek; Nguyen, Petr; Sadilek, David; Stundlova, Jana; Just, Pavel; Haddad, Charles R.; Koc, HalilWe examined the distribution of genes for major ribosomal RNAs (rDNA) on holokinetic chromosomes of 74 species belonging to 19 genera of scorpions from the family Buthidae using fluorescence in situ hybridization (FISH). Our analysis revealed differences between the two main evolutionary lineages within the family. The genera belonging to the `Buthus group', with a proposed Laurasian origin, possess one pair of rDNA mainly in an interstitial position, with the only exceptions being the terminal location found in some Hottentotta and Buthacus species, possibly as a result of chromosome fissions. All the remaining buthid 'groups' possess rDNA found strictly in a terminal position. However, the number of signals may increase from an ancestral state of one pair of rDNA loci to up to seven signals in Reddyanus ceylonensis Kovarik et al., 2016. Despite the differences in evolutionary dynamics of the rDNA clusters between the 'Buthus group' and other lineages investigated, we found a high incidence of reciprocal translocations and presence of multivalent associations during meiosis in the majority of the genera studied. These phenomena seem to be typical for the whole family Buthidae.Öğe Genetic diversity within Scorpio maurus (Scorpiones: Scorpionidae) from Turkey(Univ Oradea Publ House, 2017) Koc, Halil; Sipahi, Hulya; Yagmur, Ersen AydinIn this study, DNA sequence diversity of the mitochondrial (mt) cytochrome oxidase 1 (CO1) gene was investigated in Scorpio maurus specimens from across southeastern Turkey. Nucleotide sequences included 508 conserved sites and 126 variable sites, and the mean nucleotide variation within species was 7.9%. Intraspecific pairwise divergences ranged from 0.5% to 10.7%. Phylogenetic analysis indicated high divergence among specimens. This study is the first mtDNA sequence analysis for Turkish scorpions.Öğe Gylippus (Gylippus) erseni sp. n. (Solifugae: Gylippidae: Gylippinae), a New Solifuge Species from Turkey(Inst Zoology, Bas, 2019) Koc, Halil; Erdek, MelekA new solifuge species, Gylippus (Gylippus) erseni sp. n., is described and illustrated from Mount Karadag, Karaman Province, Turkey. It belongs to the subfamily Gylippinae (Solifugae: Gylippidae). The description is based on both males and females.Öğe Karyotypes of Southeastern Turkish Scorpions Hottentotta saulcyi and Buthacus macrocentrus (Scorpiones: Buthidae)(Istanbul University Press, 2019) Koc, Halil; Yagmur, Ersen Aydin; Šťáhlavský, FrantisekObjective: The species Hottentotta saulcyi is widely distributed from Mardin to Hakkari while the distribution of Buthacus macrocentrus is limited to the south-east of Turkey (only Şanlıurfa). The aim of this study is to analyze the cytogenetic structure of Hottentotta saulcyi and Buthacus macrocentrus. Materials and Methods: The specimens were collected during the night from Şırnak and Şanlıurfa using a UV lamp. The male Hottentotta saulcyi were collected from Şırnak and male and female Buthacus macrocentrus from Şanlıurfa. Chromosome preparations were made using cells from the male testes and the female ovariuteri of the species studied. Chromosome preparations were made using the classical spreading method. Results: The diploid chromosome number for Hottentotta saulcyi was 2n=14, and 2n=28 for Buthacus macrocentrus. Conclusion: The karyotypes of Hottentotta saulcyi and Buthacus macrocentrus have been presented for the first time. Both analyzed species have holocentric chromosomes that gradually decrease in size. Quadrivalent and hexavalent were observed during the first meiotic division in males of Buthacus macrocentrus. © Copyright 2019 by The Istanbul University Faculty of Science.Öğe Molecular cytogenetics of Androctonus scorpions: an oasis of calm in the turbulent karyotype evolution of the diverse family Buthidae(Oxford Univ Press, 2015) Sadilek, David; Nguyen, Petr; Koc, Halil; Kovarik, Frantisek; Yagmur, Ersen Aydin; Stahlavsky, FrantisekRecent cytogenetic and genomic studies suggest that morphological and molecular evolution is decoupled in the basal arachnid order Scorpiones. Extraordinary karyotype variation has been observed particularly in the family Buthidae, which is unique among scorpions for its holokinetic chromosomes. We analyzed the karyotypes of four geographically distant species of the genus AndroctonusEhrenberg, 1828 (Androctonus australis, Androctonus bourdoni, Androctonus crassicauda, Androctonus maelfaiti) (Scorpiones: Buthidae) using both classic and molecular cytogenetic methods. The mitotic complement of all species consisted of 2n=24 elements. Fluorescence in situ hybridization with a fragment of the 18S ribosomal RNA gene, a cytogenetic marker well known for its mobility, identified a single interstitial rDNA locus on the largest chromosome pair in all species examined. Our findings thus support the evolutionary stasis of the Androctonus karyotype, which is discussed with respect to current hypotheses on chromosome evolution both within and beyond the family Buthidae. Differences in karyotype dynamics between Androctonus spp. and the other buthids can help us better understand the driving forces behind their chromosome evolution and speciation.(c) 2015 The Linnean Society of London, Biological Journal of the Linnean Society, 2015, 115, 69-76.Öğe MORPHOLOGICAL COMPARISON OF THE MALLEOLI (RACQUET ORGANS) IN BITON ZEDERBAUERI AND GLUVIOPSILLA DISCOLOR (DAESIIDAE, SOLIFUGAE)(Amer Entomol Soc, 2012) Yigit, Nazife; Erdek, Melek; Koc, Halil; Melekoglu, AbdullahThe detailed morphology of the malleoli in two male solfugids, Biton zederbaueri (Werner, 1905) and Gluviopsilla discolor (Kraepelin, 1899), (Daesiidae, Solifugae) arc described and illustrated. This study was performed using stereo and scanning electron microscopy (SEM). Solifuges have five pairs of malleoli on the ventral surface of the fourth pair of legs of both sexes, as distinct from other arachnids. Malleolar sensory systems function as mechanoreceptors and chemoreceptors. A malleolus consists of two parts: a basal stalk and a triangular fan. There are epicuticular protrusions on the anterior face of each fan, and there are granular/vesicular structures on each stalk. Also, there are waved surfaces on the distal of the fan. This is the first study on the morphology of malleoli in solifuges in Turkey.Öğe New Records of Spiders (Arachnida: Araneae) from Sinop Province, Turkey, Including an Annotated List of Species(Kastamonu Univ, 2019) Etirli, Esra; Koc, Halil; Sancak, ZaferAim of study: This study was conducted to determine the biological richness of spiders from Sinop province, Turkey. Area of study: The studies were carried out on between 41 degrees 12' and 42 degrees 6' northern latitudes and 34 degrees 14' and 35 degrees 26' east longitudes within the Sinop province borders. Material and Methods: Spiders were collected from 2009 to 2015 using four collection methods: using sweep netting, tree beating, active searching and pitfall trapping. Main results: A total of 1766 specimens were collected that represented by 31 families, 133 genera and 200 species. The most diverse families collected are the Salticidae (29 spp.), Theridiidae (29 spp.) and Gnaphosidae (21 spp.). According to previous studies and this work, 31 families, 139 genera, 212 species of spiders are present in Sinop province, of which 192 species are recorded for the first time. Research highlights: Furthermore, one genus (Theridula Emerton 1882) and three species, namely Asagena meridionalis Kulczynski 1894, Theridion boesenbergi Strand 1904 and Theridula gonygaster (Simon 1873), are new records for Turkey, and are provided with illustrations. Here, we provide the first annotated checklist of the spider species of Sinop province.Öğe The first chromosome study of the genera Calchas Birula, 1899 and Neocalchas Yagmur, Soleglad, Fet & Kovarik, 2013 (Scorpiones: Iuridae)(Taylor & Francis Ltd, 2023) Yagmur, Ersen Aydin; Koc, Halil; Yesilyurt, Fatih; Stahlavsky, FrantisekThe chromosomes of two species belonging to the scorpion family Iuridae are studied for the first time. Both analysed species displayed achiasmatic meiosis and no morphologically differentiated sex chromosomes; these features are typical for all scorpions. Both species possessed monocentric chromosomes, a characteristic specific to the entire parvorder Iurida. The karyotype of Calchas nordmanni Birula, 1899 is composed of 82 chromosomes, and the diploid number of chromosomes in Neocalchas gruberi is 40. Despite this conspicuous difference in 2n chromosome morphology is mainly acrocentric. Both species possess two pairs of the 18S rDNA clusters identified by FISH. A higher number of rDNA loci may represent an ancestral state for the parvorder Iurida.Öğe The first record of karyotypes in Leiurus abdullahbayrami and Compsobuthus matthiesseni (Scorpiones: Buthidae) from Turkey(Univ Oradea Publ House, 2014) Stahlavsky, Frantisek; Koc, Halil; Yagmur, Ersen AydinThe karyotypes of Leiurus abdullahbayrami Yagmur, Koc & Kunt, 2009 and Compsobuthus matthiesseni (Birula, 1905) from Turkey are presented for the first time. Both analyzed species have achiasmatic meiosis and the same number of holocentric chromosomes (2n=22) gradually decreasing in size. During the first meiotic division we observed tetravalent in one male of L. abdullahbayrann and octovalent in one male of C. matthiesseni. These types of multivalents may increase genetic variability and indicate reciprocal translocations within these species.Öğe THE TYPE LOCALITY OF RHAGODES CAUCASICUS BIRULA, 1905, AND THE FIRST RECORD OF THE SPECIES FROM IRAN WITH REDESCRIPTION OF THE SPECIES FROM TURKEY (SOLIFUGAE: RHAGODIDAE)(Amer Entomol Soc, 2015) Koc, Halil; Moradi, Mohammed; Erdek, Melek; Yagmur, Ersen Aydin; Gharahkloo, Pooyan MoradiThe type locality of Rhagodes caucasicus Birula, 1905 is corrected and identified as Tuzluca (Kul'p), Igdir Province, eastern Turkey instead of Armenia. A new record for the Iranian solpugid fauna is also reported: Rhagodes caucasicus Birula, 1905 from Zanjan Province, northwestern Iran. The redescription of new material of Rhagodes caucasicus from Turkey is included.Öğe TWO NEW RECORDS FOR TURKISH HARVESTMEN FAUNA (ARACHNIDA: OPILIONES)(Amer Entomol Soc, 2015) Kurt, Kemal; Koc, Halil; Yagmur, Ersen AydinTwo harvestmen species, Mediostoma haasi (Roewer, 1953) and Phalangium armatum Snegovaya, 2005, are recorded in Turkey for the first time. Their description, characteristic features, distributions, measurements and photographs of genitalia and general habitus of both sexes are presented.
